Help with Pilea Peperomioides

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My Pilea peperomioides is doing pretty well overall, but sometimes I notice some leaves yellowing. It seems to do especially poorly in the winter and usually recovers in the summer. I notice that the leaves start out normal, then they look like the ones in the picture, and then they turn completely yellow, then brown and fall off. Still growing quite a bit during the summer though.

See the photos for details. I have the plant in a sunroom (no windows on the ceiling, but the south wall is a 1 big window and the larger west-facing side is all windows. No windows on the north side). Lots of bright indirect light all year round, but I live in the PNW, so there are lots of clouds and not much light in the winter. Some direct light when the sun is shining through the windows. Temps get up to 90 degrees in the summer (but good ventilation with breezes blowing through), down to 40s in the winter. I use a soil moisture monitor and water whenever the plant is totally dry.

Any advice?
 

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